Browsing by Author "Groffman, Peter M."
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- Climate and lawn management interact to control C-4 plant distribution in residential lawns across seven US citiesTrammell, Tara L. E.; Pataki, Diane E.; Still, Christopher J.; Ehleringer, James R.; Avolio, Meghan L.; Bettez, Neil; Cavender-Bares, Jeannine; Groffman, Peter M.; Grove, J. Morgan; Hall, Sharon J.; Heffernan, James B.; Hobbie, Sarah E.; Larson, Kelli L.; Morse, Jennifer L.; Neill, Christopher; Nelson, Kristen C.; O'Neil-Dunne, Jarlath P.M.; Pearse, William D.; Chowdhury, Rinku Roy; Steele, Meredith K.; Wheeler, Megan M. (2019-06)In natural grasslands, C-4 plant dominance increases with growing season temperatures and reflects distinct differences in plant growth rates and water use efficiencies of C-3 vs. C-4 photosynthetic pathways. However, in lawns, management decisions influence interactions between planted turfgrass and weed species, leading to some uncertainty about the degree of human vs. climatic controls on lawn species distributions. We measured herbaceous plant carbon isotope ratios (delta C-13, index of C-3/C-4 relative abundance) and C-4 cover in residential lawns across seven U.S. cities to determine how climate, lawn plant management, or interactions between climate and plant management influenced C-4 lawn cover. We also calculated theoretical C-4 carbon gain predicted by a plant physiological model as an index of expected C-4 cover due to growing season climatic conditions in each city. Contrary to theoretical predictions, plant delta C-13 and C-4 cover in urban lawns were more strongly related to mean annual temperature than to growing season temperature. Wintertime temperatures influenced the distribution of C-4 lawn turf plants, contrary to natural ecosystems where growing season temperatures primarily drive C-4 distributions. C-4 cover in lawns was greatest in the three warmest cities, due to an interaction between climate and homeowner plant management (e.g., planting C-4 turf species) in these cities. The proportion of C-4 lawn species was similar to the proportion of C-4 species in the regional grass flora. However, the majority of C-4 species were nonnative turf grasses, and not of regional origin. While temperature was a strong control on lawn species composition across the United States, cities differed as to whether these patterns were driven by cultivated lawn grasses vs. weedy species. In some cities, biotic interactions with weedy plants appeared to dominate, while in other cities, C-4 plants were predominantly imported and cultivated. Elevated CO2 and temperature in cities can influence C-3/C-4 competitive outcomes; however, this study provides evidence that climate and plant management dynamics influence biogeography and ecology of C-3/C-4 plants in lawns. Their differing water and nutrient use efficiency may have substantial impacts on carbon, water, energy, and nutrient budgets across cities.
- Connectivity: insights from the US Long Term Ecological Research NetworkIwaniec, David M.; Gooseff, Michael N.; Suding, Katharine N.; Samuel Johnson, David; Reed, Daniel C.; Peters, Debra P. C.; Adams, Byron J.; Barrett, John E.; Bestelmeyer, Brandon T.; Castorani, Max C. N.; Cook, Elizabeth M.; Davidson, Melissa J.; Groffman, Peter M.; Hanan, Niall P.; Huenneke, Laura F.; Johnson, Pieter T. J.; McKnight, Diane M.; Miller, Robert J.; Okin, Gregory S.; Preston, Daniel L.; Rassweiler, Andrew; Ray, Chris; Sala, Osvaldo E.; Schooley, Robert L.; Seastedt, Timothy; Spasojevic, Marko J.; Vivoni, Enrique R. (2021-05)Ecosystems across the United States are changing in complex and surprising ways. Ongoing demand for critical ecosystem services requires an understanding of the populations and communities in these ecosystems in the future. This paper represents a synthesis effort of the U.S. National Science Foundation-funded Long-Term Ecological Research (LTER) network addressing the core research area of "populations and communities." The objective of this effort was to show the importance of long-term data collection and experiments for addressing the hardest questions in scientific ecology that have significant implications for environmental policy and management. Each LTER site developed at least one compelling case study about what their site could look like in 50-100 yr as human and environmental drivers influencing specific ecosystems change. As the case studies were prepared, five themes emerged, and the studies were grouped into papers in this LTER Futures Special Feature addressing state change, connectivity, resilience, time lags, and cascading effects. This paper addresses the "connectivity" theme and has examples from the Phoenix (urban), Niwot Ridge (alpine tundra), McMurdo Dry Valleys (polar desert), Plum Island (coastal), Santa Barbara Coastal (coastal), and Jornada (arid grassland and shrubland) sites. Connectivity has multiple dimensions, ranging from multi-scalar interactions in space to complex interactions over time that govern the transport of materials and the distribution and movement of organisms. The case studies presented here range widely, showing how land-use legacies interact with climate to alter the structure and function of arid ecosystems and flows of resources and organisms in Antarctic polar desert, alpine, urban, and coastal marine ecosystems. Long-term ecological research demonstrates that connectivity can, in some circumstances, sustain valuable ecosystem functions, such as the persistence of foundation species and their associated biodiversity or, it can be an agent of state change, as when it increases wind and water erosion. Increased connectivity due to warming can also lead to species range expansions or contractions and the introduction of undesirable species. Continued long-term studies are essential for addressing the complexities of connectivity. The diversity of ecosystems within the LTER network is a strong platform for these studies.
- Continental-scale homogenization of residential lawn plant communitiesWheeler, Megan M.; Neill, Christopher; Groffman, Peter M.; Avolio, Meghan L.; Bettez, Neil; Cavender-Bares, Jeannine; Chowdhury, Rinku Roy; Darling, Lindsay E.; Grove, J. Morgan; Hall, Sharon J.; Heffernan, James B.; Hobbie, Sarah E.; Larson, Kelli L.; Morse, Jennifer L.; Nelson, Kristen C.; Ogden, Laura A.; O'Neil-Dunne, Jarlath P.M.; Pataki, Diane E.; Polsky, Colin; Steele, Meredith K.; Trammell, Tara L. E. (2017-09)Residential lawns are highly managed ecosystems that occur in urbanized landscapes across the United States. Because they are ubiquitous, lawns are good systems in which to study the potential homogenizing effects of urban land use and management together with the continental-scale effects of climate on ecosystem structure and functioning. We hypothesized that similar homeowner preferences and management in residential areas across the United States would lead to low plant species diversity in lawns and relatively homogeneous vegetation across broad geographical regions. We also hypothesized that lawn plant species richness would increase with regional temperature and precipitation due to the presence of spontaneous, weedy vegetation, but would decrease with household income and fertilizer use. To test these predictions, we compared plant species composition and richness in residential lawns in seven U.S. metropolitan regions. We also compared species composition in lawns with understory vegetation in minimally-managed reference areas in each city. As expected, the composition of cultivated turfgrasses was more similar among lawns than among reference areas, but this pattern also held among spontaneous species. Plant species richness and diversity varied more among lawns than among reference areas, and more diverse lawns occurred in metropolitan areas with higher precipitation. Native forb diversity increased with precipitation and decreased with income, driving overall lawn diversity trends with these predictors as well. Our results showed that both management and regional climate shaped lawn species composition, but the overall homogeneity of species regardless of regional context strongly suggested that management was a more important driver.
- Convergence of microclimate in residential landscapes across diverse cities in the United StatesHall, Sharon J.; Learned, Jennifer; Ruddell, Benjamin L.; Larson, K. L.; Cavender-Bares, Jeannine; Bettez, Neil; Groffman, Peter M.; Grove, J. Morgan; Heffernan, James B.; Hobbie, Sarah E.; Morse, Jennifer L.; Neill, Christopher; Nelson, Kristen C.; O'Neil-Dunne, Jarlath P.M.; Ogden, Laura A.; Pataki, Diane E.; Pearse, William D.; Polsky, Colin; Chowdhury, Rinku Roy; Steele, Meredith K.; Trammell, Tara L. E. (2016-01)The urban heat island (UHI) is a well-documented pattern of warming in cities relative to rural areas. Most UHI research utilizes remote sensing methods at large scales, or climate sensors in single cities surrounded by standardized land cover. Relatively few studies have explored continental-scale climatic patterns within common urban microenvironments such as residential landscapes that may affect human comfort. We tested the urban homogenization hypothesis which states that structure and function in cities exhibit ecological "sameness" across diverse regions relative to the native ecosystems they replaced. We deployed portable micrometeorological sensors to compare air temperature and humidity in residential yards and native landscapes across six U.S. cities that span a range of climates (Phoenix, AZ; Los Angeles, CA; Minneapolis-St. Paul, MN; Boston, MA; Baltimore, MD; and Miami, FL). Microclimate in residential ecosystems was more similar among cities than among native ecosystems, particularly during the calm morning hours. Maximum regional actual evapotranspiration (AET) was related to the morning residential microclimate effect. Residential yards in cities with maximum AET < 50-65 cm/year (Phoenix and Los Angeles) were generally cooler and more humid than nearby native shrublands during summer mornings, while yards in cities above this threshold were generally warmer (Baltimore and Miami) and drier (Miami) than native forests. On average, temperature and absolute humidity were similar to 6 % less variable among residential ecosystems than among native ecosystems from diverse regions. These data suggest that common residential land cover and structural characteristics lead to microclimatic convergence across diverse regions at the continental scale.
- Drivers of plant species richness and phylogenetic composition in urban yards at the continental scaleCubino, Josep Padullés; Cavender-Bares, Jeannine; Hobbie, Sarah E.; Pataki, Diane E.; Avolio, Meghan L.; Darling, Lindsay E.; Larson, Kelli L.; Hall, Sharon J.; Groffman, Peter M.; Trammell, Tara L. E.; Steele, Meredith K.; Grove, J. Morgan; Neill, Christopher (2018-11-23)Context As urban areas increase in extent globally, domestic yards play an increasingly important role as potential contributors to ecosystem services and well-being. These benefits largely depend on the plant species richness and composition of yards. Objectives We aim to determine the factors that drive plant species richness and phylogenetic composition of cultivated and spontaneous flora in urban yards at the continental scale, and how these potential drivers interact. Methods We analyzed plant species richness and phylogenetic composition of cultivated and spontaneous flora of 117 private yards from six major metropolitan areas in the US. Yard plant species richness and phylogenetic composition were expressed as a function of biophysical and socioeconomic variables and yard characteristics using linear mixed-effects models and spatially explicit structural equation modeling. Results Extreme temperatures largely determined yard species richness and phylogenetic composition at the continental scale. Precipitation positively predicted spontaneous richness but negatively predicted cultivated richness. Only the phylogenetic composition of the spontaneous flora was associated with precipitation. The effect of lower temperatures and precipitation on all yard diversity parameters was partly mediated by yard area. Among various socioeconomic variables, only education level showed a significant effect on cultivated phylogenetic composition. Conclusions Our results support the hypothesis that irrigation compensates for precipitation in driving cultivated yard plant diversity at the continental scale. Socioeconomic variables among middle and upper class families have no apparent influence on yard diversity. These findings inform the adaptation of US urban vegetation in cities in the face of global change.
- Ecological homogenization of soil properties in the American residential macrosystemRyan, Christopher D.; Groffman, Peter M.; Grove, J. Morgan; Hall, Sharon J.; Heffernan, James B.; Hobbie, Sarah E.; Locke, Dexter H.; Morse, Jennifer L.; Neill, Christopher; Nelson, Kristen C.; O'Neil-Dunne, Jarlath; Chowdhury, Rinku Roy; Steele, Meredith K.; Trammell, Tara L. E. (Wiley, 2022-09)The conversion of native ecosystems to residential ecosystems dominated by lawns has been a prevailing land-use change in the United States over the past 70 years. Similar development patterns and management of residential ecosystems cause many characteristics of residential ecosystems to be more similar to each other across broad continental gradients than that of former native ecosystems. For instance, similar lawn management by irrigation and fertilizer applications has the potential to influence soil carbon (C) and nitrogen (N) pools and processes. We evaluated the mean and variability of total soil C and N stocks, potential net N mineralization and nitrification, soil nitrite (NO2-)/nitrate (NO3-) and ammonium (NH4+) pools, microbial biomass C and N content, microbial respiration, bulk density, soil pH, and moisture content in residential lawns and native ecosystems in six metropolitan areas across a broad climatic gradient in the United States: Baltimore, MD (BAL); Boston, MA (BOS); Los Angeles, CA (LAX); Miami, FL (MIA); Minneapolis-St. Paul, MN (MSP); and Phoenix, AZ (PHX). We observed evidence of higher N cycling in lawn soils, including significant increases in soil NO2-/NO3-, microbial N pools, and potential net nitrification, and significant decreases in NH4+ pools. Self-reported yard fertilizer application in the previous year was linked with increased NO2-/ NO3- content and decreases in total soil N and C content. Self-reported irrigation in the previous year was associated with decreases in potential net mineralization and potential net nitrification and with increases in bulk density and pH. Residential topsoil had higher total soil C than native topsoil, and microbial biomass C was markedly higher in residential topsoil in the two driest cities (LAX and PHX). Coefficients of variation for most biogeochemical metrics were higher in native soils than in residential soils across all cities, suggesting that residential development homogenizes soil properties and processes at the continental scale.
- Ecosystem services in managing residential landscapes: priorities, value dimensions, and cross-regional patternsLarson, K. L.; Nelson, Kristen C.; Samples, S. R.; Hall, Sharon J.; Bettez, Neil; Cavender-Bares, Jeannine; Groffman, Peter M.; Grove, J. Morgan; Heffernan, James B.; Hobbie, Sarah E.; Learned, Jennifer; Morse, Jennifer L.; Neill, Christopher; Ogden, Laura A.; O'Neil-Dunne, Jarlath P.M.; Pataki, Diane E.; Polsky, Colin; Chowdhury, Rinku Roy; Steele, Meredith K.; Trammell, Tara L. E. (2016-03)Although ecosystem services have been intensively examined in certain domains (e.g., forests and wetlands), little research has assessed ecosystem services for the most dominant landscape type in urban ecosystems-namely, residential yards. In this paper, we report findings of a cross-site survey of homeowners in six U.S. cities to 1) examine how residents subjectively value various ecosystem services, 2) explore distinctive dimensions of those values, and 3) test the urban homogenization hypothesis. This hypothesis posits that urbanization leads to similarities in the social-ecological dynamics across cities in diverse biomes. By extension, the thesis suggests that residents' ecosystem service priorities for residential landscapes will be similar regardless of whether residents live in the humid East or the arid West, or the warm South or the cold North. Results underscored that cultural services were of utmost importance, particularly anthropocentric values including aesthetics, low-maintenance, and personal enjoyment. Using factor analyses, distinctive dimensions of residents' values were found to partially align with the Millennium Ecosystem Assessment's categories (provisioning, regulating, supporting, and cultural). Finally, residents' ecosystem service priorities exhibited significant homogenization across regions. In particular, the traditional lawn aesthetic (neat, green, weed-free yards) was similarly important across residents of diverse U.S. cities. Only a few exceptions were found across different environmental and social contexts; for example, cooling effects were more important in the warm South, where residents also valued aesthetics more than those in the North, where low-maintenance yards were a greater priority.
- Homogenization of plant diversity, composition, and structure in North American urban yardsPearse, William D.; Cavender-Bares, Jeannine; Hobbie, Sarah E.; Avolio, Meghan L.; Bettez, Neil; Chowdhury, Rinku Roy; Darling, Lindsay E.; Groffman, Peter M.; Grove, J. Morgan; Hall, Sharon J.; Heffernan, James B.; Learned, Jennifer; Neill, Christopher; Nelson, Kristen C.; Pataki, Diane E.; Ruddell, Benjamin L.; Steele, Meredith K.; Trammell, Tara L. E. (Ecological Society of America, 2018-02)Urban ecosystems are widely hypothesized to be more ecologically homogeneous than natural ecosystems. We argue that urban plant communities assemble from a complex mix of horticultural and regional species pools, and evaluate the homogenization hypothesis by comparing cultivated and spontaneously occurring urban vegetation to natural area vegetation across seven major U.S. cities. There was limited support for homogenization of urban diversity, as the cultivated and spontaneous yard flora had greater numbers of species than natural areas, and cultivated phylogenetic diversity was also greater. However, urban yards showed evidence of homogenization of composition and structure. Yards were compositionally more similar across regions than were natural areas, and tree density was less variable in yards than in comparable natural areas. This homogenization of biodiversity likely reflects similar horticultural source pools, homeowner preferences, and management practices across U.S. cities.
- Isotopic signals of summer denitrification in a northern hardwood forested catchmentWexler, S. K.; Goodale, Christine L.; McGuire, Kevin J.; Bailey, Scott W.; Groffman, Peter M. (National Academy of Sciences, 2014-11-18)Despite decades of measurements, the nitrogen balance of temperate forest catchments remains poorly understood. Atmospheric nitrogen deposition often greatly exceeds streamwater nitrogen losses; the fate of the remaining nitrogen is highly uncertain. Gaseous losses of nitrogen to denitrification are especially poorly documented and are often ignored. Here, we provide isotopic evidence (δ15NNO3 and δ18ONO3) from shallow groundwater at the Hubbard Brook Experimental Forest indicating extensive denitrification during midsummer, when transient, perched patches of saturation developed in hillslopes, with poor hydrological connectivity to the stream, while streamwater showed no isotopic evidence of denitrification. During small rain events, precipitation directly contributed up to 34% of streamwater nitrate, which was otherwise produced by nitrification. Together, these measurements reveal the importance of denitrification in hydrologically disconnected patches of shallow groundwater during midsummer as largely overlooked control points for nitrogen loss from temperate forest catchments.
- Searching for biogeochemical hot spots in three dimensions: Soil C and N cycling in hydropedologic settings in a northern hardwood forestMorse, J. L.; Werner, S. F.; Gillin, C. P.; Goodale, Christine L.; Bailey, Scott W.; McGuire, Kevin J.; Groffman, Peter M. (American Geophysical Union, 2014-08-01)Understanding and predicting the extent, location, and function of biogeochemical hot spots at the watershed scale is a frontier in environmental science. We applied a hydropedologic approach to identify (1) biogeochemical differences among morphologically distinct hydropedologic settings and (2) hot spots of microbial carbon (C) and nitrogen (N) cycling activity in a northern hardwood forest in Hubbard Brook Experimental Forest, New Hampshire, USA. We assessed variables related to C and N cycling in spodic hydropedologic settings (typical podzols, bimodal podzols, and Bh podzols) and groundwater seeps during August 2010. We found that soil horizons (Oi/Oe, Oa/A, and B) differed significantly for most variables. B horizons (>10cm) accounted for 71% (±11%) of C pools and 62%(±10%) ofmicrobial biomass C in the sampled soil profile, whereas the surface horizons (Oi/Oe and Oa/A; 0–10cm) were dominant zones for N-cycle-related variables. Watershed-wide estimates of C and N cycling were higher by 34 to 43% (±17–19%) when rates, horizon thickness, and areal extent of each hydropedologic setting were incorporated, versus conventionally calculated estimates for typical podzols that included only the top 10cm of mineral soil. Despite the variation in profile development in typical, bimodal, and Bh podzols, we did not detect significant differences in C and N cycling among them. Across all soil horizons and hydropedologic settings, we found strong links between biogeochemical cycling and soil C, suggesting that the accumulation of C in soils may be a robust indicator of microbial C and N cycling capacity in the landscape.
- Sediment chemistry of urban stormwater ponds and controls on denitrificationBlaszczak, Joanna R.; Steele, Meredith K.; Badgley, Brian D.; Heffernan, James B.; Hobbie, Sarah E.; Morse, Jennifer L.; Rivers, Erin N.; Hall, Sharon J.; Neill, Christopher; Pataki, Diane E.; Groffman, Peter M.; Bernhardt, Emily S. (Ecological Society of America, 2018-06)Stormwater ponds and retention basins are ubiquitous features throughout urban landscapes. These ponds are potentially important control points for nitrogen (N) removal from surface water bodies via denitrification. However, there are possible trade-offs to this water quality benefit if high N and contaminant concentrations in stormwater pond sediments decrease the complete reduction of nitrous oxide (N2O), a potent greenhouse gas, to dinitrogen (N-2) during denitrification. This may occur through decreasing the abundance or efficiency of denitrifiers capable of producing the N2O reductase enzyme. We predicted that ponds draining increasingly urbanized landscapes would have higher N and metal concentrations in their sediments, and thereby greater N2O yields. We measured potential denitrification rates, N2O reductase (nosZ) gene frequencies, as well as sediment and pore water chemistry in 64 ponds distributed across eight U.S. cities. We found almost no correlation between the proportion of urban land cover surrounding ponds and the nutrient and contaminant concentrations in the stormwater pond sediments within or across all cities. Regression analysis revealed that the proportion of potential N-2 and N2O production that could be explained was under different environmental controls. Our survey raises many new questions about why N fluxes and transformations vary so widely both within and across urban environments, but also allays the concern that elevated metal concentrations in urban stormwater ponds will increase N2O emissions. Urban stormwater ponds are unlikely to be a problematic source of N2O to the atmosphere, no matter their denitrification potential.
- Urban soil carbon and nitrogen converge at a continental scaleTrammell, Tara L. E.; Pataki, Diane E.; Pouyat, Richard, V; Groffman, Peter M.; Rosier, Carl; Bettez, Neil; Cavender-Bares, Jeannine; Grove, J. Morgan; Hall, Sharon J.; Heffernan, James B.; Hobbie, Sarah E.; Morse, Jennifer L.; Neill, Christopher; Steele, Meredith K. (2020-05)In urban areas, anthropogenic drivers of ecosystem structure and function are thought to predominate over larger-scale biophysical drivers. Residential yards are influenced by individual homeowner preferences and actions, and these factors are hypothesized to converge yard structure across broad scales. We examined soil total C and total delta C-13, organic C and organic delta C-13, total N, and delta N-15 in residential yards and corresponding reference ecosystems in six cities across the United States that span major climates and ecological biomes (Baltimore, Maryland; Boston, Massachusetts; Los Angeles, California; Miami, Florida; Minneapolis-St. Paul, Minnesota; and Phoenix, Arizona). Across the cities, we found soil C and N concentrations and soil delta N-15 were less variable in residential yards compared to reference sites supporting the hypothesis that soil C, N, and delta N-15 converge across these cities. Increases in organic soil C, soil N, and soil delta N-15 across urban, suburban, and rural residential yards in several cities supported the hypothesis that soils responded similarly to altered resource inputs across cities, contributing to convergence of soil C and N in yards compared to natural systems. Soil C and N dynamics in residential yards showed evidence of increasing C and N inputs to urban soils or dampened decomposition rates over time that are influenced by climate and/or housing age across the cities. In the warmest cities (Los Angeles, Miami, Phoenix), greater organic soil C and higher soil delta C-13 in yards compared to reference sites reflected the greater proportion of C-4 plants in these yards. In the two warm arid cities (Los Angeles, Phoenix), total soil delta C-13 increased and organic soil delta C-13 decreased with increasing home age indicating greater inorganic C in the yards around newer homes. In general, soil organic C and delta C-13, soil N, and soil delta N-15 increased with increasing home age suggesting increased soil C and N cycling rates and associated C-12 and N-14 losses over time control yard soil C and N dynamics. This study provides evidence that conversion of native reference ecosystems to residential areas results in convergence of soil C and N at a continental scale. The mechanisms underlying these effects are complex and vary spatially and temporally.
- Watershed studies at the Hubbard Brook Experimental Forest: Building on a long legacy of research with new approaches and sources of dataCampbell, John L.; Rustad, Lindsey E.; Bailey, Scott W.; Bernhardt, Emily S.; Driscoll, Charles T.; Green, Mark B.; Groffman, Peter M.; Lovett, Gary M.; McDowell, William H.; McGuire, Kevin J.; Rosi, Emma J. (2021-01)The Hubbard Brook Experimental Forest (HBEF) was established in 1955 by the U.S. Department of Agriculture, Forest Service out of concerns about the effects of logging increasing flooding and erosion. To address this issue, within the HBEF hydrological and micrometeorological monitoring was initiated in small watersheds designated for harvesting experiments. The Hubbard Brook Ecosystem Study (HBES) originated in 1963, with the idea of using the small watershed approach to study element fluxes and cycling and the response of forest ecosystems to disturbances, such as forest management practices and air pollution. Early evidence of acid rain was documented at the HBEF and research by scientists at the site helped shape acid rain mitigation policies. New lines of investigation at the HBEF have built on the long legacy of watershed research resulting in a shift from comparing inputs and outputs and quantifying pools and fluxes to a more mechanistic understanding of ecosystem processes within watersheds. For example, hydropedological studies have shed light on linkages between hydrologic flow paths and soil development that provide valuable perspective for managing forests and understanding stream water quality. New high frequency in situ stream chemistry sensors are providing insights about extreme events and diurnal patterns that were indiscernible with traditional weekly sampling. Additionally, tools are being developed for visual and auditory data exploration and discovery by a broad audience. Given the unprecedented environmental change that is occurring, data from the small watersheds at the HBEF are more relevant now than ever and will continue to serve as a basis for sound environmental decision-making.